ATH-PA

SEC source: 0001127602-19-001369 · form4.xml · nonDerivative · entry 4

WHEELER WILLIAM J · CIK 0001273281 · President

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 398,235

F5 · shares: This reported transaction represents shares withheld to satisfy tax withholding obligations in connection with the settlement of vested RSUs that settled in Class A common shares.

SEC source: 0001127602-19-001369 · form4.xml · nonDerivative · entry 3

WHEELER WILLIAM J · CIK 0001273281 · President

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 401,283

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F4 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001369 · form4.xml · nonDerivative · entry 2

WHEELER WILLIAM J · CIK 0001273281 · President

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 398,682

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F4 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001369 · form4.xml · nonDerivative · entry 1

WHEELER WILLIAM J · CIK 0001273281 · President

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 396,243

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F4 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001381 · form4.xml · derivative · entry 7

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 3,469

Underlying: Class A Common Shares · 1,734 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F3 · derivative.exercise_price: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date.

F4 · derivative.exercise_price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F3 · derivative.exercise_date: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date.

F3 · derivative.expiration_date: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date.

SEC source: 0001127602-19-001381 · form4.xml · derivative · entry 6

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 1,627

Underlying: Class A Common Shares · 1,626 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F2 · derivative.exercise_price: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2017 vesting start date. An additional 9,757 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2017 to December 31, 2019.

F4 · derivative.exercise_price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F2 · derivative.exercise_date: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2017 vesting start date. An additional 9,757 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2017 to December 31, 2019.

F2 · derivative.expiration_date: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2017 vesting start date. An additional 9,757 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2017 to December 31, 2019.

SEC source: 0001127602-19-001381 · form4.xml · derivative · entry 5

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 0

Underlying: Class A Common Shares · 1,841 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F1 · derivative.exercise_price: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2016 vesting start date. An additional 11,046 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2016 to December 31, 2018.

F4 · derivative.exercise_price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F1 · derivative.exercise_date: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2016 vesting start date. An additional 11,046 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2016 to December 31, 2018.

F1 · derivative.expiration_date: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2016 vesting start date. An additional 11,046 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2016 to December 31, 2018.

SEC source: 0001127602-19-001381 · form4.xml · nonDerivative · entry 4

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 86,833

F5 · shares: This reported transaction represents shares withheld to satisfy tax withholding obligations in connection with the settlement of vested RSUs that settled in Class A common shares.

SEC source: 0001127602-19-001381 · form4.xml · nonDerivative · entry 3

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 88,902

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F4 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001381 · form4.xml · nonDerivative · entry 2

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 87,168

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F4 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001381 · form4.xml · nonDerivative · entry 1

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 85,542

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F4 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F4 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001387 · form4.xml · derivative · entry 5

Rhodes John M · CIK 0001657130 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 2,082

Underlying: Class A Common Shares · 1,040 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F2 · derivative.exercise_price: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date.

F3 · derivative.exercise_price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F3 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F2 · derivative.exercise_date: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date.

F2 · derivative.expiration_date: Each RSU represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date.

SEC source: 0001127602-19-001387 · form4.xml · derivative · entry 4

Rhodes John M · CIK 0001657130 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 976

Underlying: Class A Common Shares · 976 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F1 · derivative.exercise_price: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2017 vesting start date. An additional 5,854 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2017 to December 31, 2019.

F3 · derivative.exercise_price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F3 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F1 · derivative.exercise_date: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2017 vesting start date. An additional 5,854 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2017 to December 31, 2019.

F1 · derivative.expiration_date: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2017 vesting start date. An additional 5,854 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2017 to December 31, 2019.

SEC source: 0001127602-19-001387 · form4.xml · nonDerivative · entry 3

Rhodes John M · CIK 0001657130 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 15,273

F4 · shares: This reported transaction represents shares withheld to satisfy tax withholding obligations in connection with the settlement of vested RSUs that settled in Class A common shares.

SEC source: 0001127602-19-001387 · form4.xml · nonDerivative · entry 2

Rhodes John M · CIK 0001657130 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 16,084

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F3 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F3 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-19-001387 · form4.xml · nonDerivative · entry 1

Rhodes John M · CIK 0001657130 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 15,044

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F3 · shares: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

F3 · price: This reported transaction represents vested RSUs that have been settled in Class A common shares. Vested RSUs are settled in Class A common shares on a one-for-one basis.

SEC source: 0001127602-18-033221 · form4.xml · derivative · entry 1

Klein Martin P · CIK 0001518052 · Executive VP and CFO

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 195,878.11

Underlying: Class A Common Shares · 7,954.34 shares

Expires not disclosed

F1 · derivative.exercise_price: These Class M-4 Prime common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F2 · shares: This reported transaction represents shares withheld to satisfy tax withholding obligations in connection with the vesting of Class M-4 Prime common shares. The price of derivative security represents the per share value of the Class M-4 Prime common shares used to calculate the number of shares withheld.

F2 · price: This reported transaction represents shares withheld to satisfy tax withholding obligations in connection with the vesting of Class M-4 Prime common shares. The price of derivative security represents the per share value of the Class M-4 Prime common shares used to calculate the number of shares withheld.

F1 · derivative.exercise_date: These Class M-4 Prime common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F3 · derivative.exercise_date: One-third of the Class M-4 Prime common shares vest based on time, with such shares vesting ratably on each of the first five anniversaries of the November 15, 2015 vesting start date; one-third vest upon the trading price of the Class A common shares attaining a per-share volume weighted average closing trading price of $50 or more during any 120-day period (or upon a sale or change in control in which Class A common shares are valued at $50 or more); and one-third vest upon the trading price of the Class A common shares attaining a per-share volume weighted average closing trading price of $70 or more during any 120-day period (or upon a sale or change in control in which Class A common shares are valued at $70 or more). Any remaining unvested Class M-4 Prime common shares expire November 15, 2025.

F1 · derivative.expiration_date: These Class M-4 Prime common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F3 · derivative.expiration_date: One-third of the Class M-4 Prime common shares vest based on time, with such shares vesting ratably on each of the first five anniversaries of the November 15, 2015 vesting start date; one-third vest upon the trading price of the Class A common shares attaining a per-share volume weighted average closing trading price of $50 or more during any 120-day period (or upon a sale or change in control in which Class A common shares are valued at $50 or more); and one-third vest upon the trading price of the Class A common shares attaining a per-share volume weighted average closing trading price of $70 or more during any 120-day period (or upon a sale or change in control in which Class A common shares are valued at $70 or more). Any remaining unvested Class M-4 Prime common shares expire November 15, 2025.

SEC source: 0001127602-18-030072 · form4.xml · derivative · entry 8

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by The Anne E Fowler Foundation, Inc.

Disclosed balance after: 19,000

Underlying: Class A Common Shares · 91 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F5 · derivative.exercise_price: These Class M-1 common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F6 · derivative.exercise_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

F6 · derivative.expiration_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

F2 · ownership.nature: The Reporting Person disclaims beneficial ownership of the shares held by The Anne E. Fowler Foundation, Inc., and this report should not be deemed an admission that the Reporting Person is the beneficial owner of such shares for purposes of Section 16 or for any other purpose.

SEC source: 0001127602-18-030072 · form4.xml · derivative · entry 7

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Gillis Family Trust U/a/d 12/30/14

Disclosed balance after: 376,000

Underlying: Class A Common Shares · 364 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F5 · derivative.exercise_price: These Class M-1 common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F6 · derivative.exercise_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

F6 · derivative.expiration_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

SEC source: 0001127602-18-030072 · form4.xml · nonDerivative · entry 5

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by The Anne E Fowler Foundation, Inc.

Disclosed balance after: 0

F3 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person, the Gillis Family Trust and The Anne E. Fowler Foundation, Inc. on March 15, 2018.

F4 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.21 inclusive. The Reporting Person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (4) to this Form 4.

F2 · ownership.nature: The Reporting Person disclaims beneficial ownership of the shares held by The Anne E. Fowler Foundation, Inc., and this report should not be deemed an admission that the Reporting Person is the beneficial owner of such shares for purposes of Section 16 or for any other purpose.

SEC source: 0001127602-18-030072 · form4.xml · nonDerivative · entry 4

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by Gillis Family Trust U/a/d 12/30/14

Disclosed balance after: 0

F3 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person, the Gillis Family Trust and The Anne E. Fowler Foundation, Inc. on March 15, 2018.

F4 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.21 inclusive. The Reporting Person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (4) to this Form 4.

SEC source: 0001127602-18-030072 · form4.xml · nonDerivative · entry 3

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 218,638

F3 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person, the Gillis Family Trust and The Anne E. Fowler Foundation, Inc. on March 15, 2018.

F4 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.21 inclusive. The Reporting Person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (4) to this Form 4.

SEC source: 0001127602-18-030072 · form4.xml · nonDerivative · entry 2

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by The Anne E Fowler Foundation, Inc.

Disclosed balance after: 91

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F2 · ownership.nature: The Reporting Person disclaims beneficial ownership of the shares held by The Anne E. Fowler Foundation, Inc., and this report should not be deemed an admission that the Reporting Person is the beneficial owner of such shares for purposes of Section 16 or for any other purpose.

SEC source: 0001127602-18-030072 · form4.xml · nonDerivative · entry 1

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by Gillis Family Trust U/a/d 12/30/14

Disclosed balance after: 364

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F1 · ownership.nature: Anne Marie Heiser is trustee of the Gillis Family Trust u/a/d 12/30/14.

SEC source: 0001127602-18-028455 · form4.xml · derivative · entry 11

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by The Anne E Fowler Foundation, Inc.

Disclosed balance after: 19,091

Underlying: Class A Common Shares · 5,909 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F6 · derivative.exercise_price: These Class M-1 common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F8 · derivative.exercise_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

F8 · derivative.expiration_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

F2 · ownership.nature: The Reporting Person disclaims beneficial ownership of the shares held by The Anne E. Fowler Foundation, Inc., and this report should not be deemed an admission that the Reporting Person is the beneficial owner of such shares for purposes of Section 16 or for any other purpose.

SEC source: 0001127602-18-028455 · form4.xml · derivative · entry 10

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Gillis Family Trust U/a/d 12/30/14

Disclosed balance after: 376,364

Underlying: Class A Common Shares · 23,636 shares

Expires not disclosed

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F6 · derivative.exercise_price: These Class M-1 common shares are exchangeable into Class A common shares, on a one-to-one basis, once vested and upon payment of the conversion price (which can be paid in cash or shares).

F8 · derivative.exercise_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

F8 · derivative.expiration_date: These Class M-1 common shares are fully vested. These vested Class M-1 common shares can be converted at the holder's election into Class A common shares at any time.

SEC source: 0001127602-18-028455 · form4.xml · nonDerivative · entry 6

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by The Anne E Fowler Foundation, Inc.

Disclosed balance after: 0

F3 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person, the Gillis Family Trust and The Anne E. Fowler Foundation, Inc. on March 15, 2018.

F4 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $51.34 to $52.29, inclusive. The Reporting Person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (4) to this Form 4.

F2 · ownership.nature: The Reporting Person disclaims beneficial ownership of the shares held by The Anne E. Fowler Foundation, Inc., and this report should not be deemed an admission that the Reporting Person is the beneficial owner of such shares for purposes of Section 16 or for any other purpose.

SEC source: 0001127602-18-028455 · form4.xml · nonDerivative · entry 5

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by Gillis Family Trust U/a/d 12/30/14

Disclosed balance after: 0

F3 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person, the Gillis Family Trust and The Anne E. Fowler Foundation, Inc. on March 15, 2018.

F5 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $51.34 to $52.16, inclusive. The Reporting Person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (5) to this Form 4.

SEC source: 0001127602-18-028455 · form4.xml · nonDerivative · entry 4

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 219,183

F3 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person, the Gillis Family Trust and The Anne E. Fowler Foundation, Inc. on March 15, 2018.

F4 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $51.34 to $52.29, inclusive. The Reporting Person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (4) to this Form 4.

SEC source: 0001127602-18-028455 · form4.xml · nonDerivative · entry 3

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by The Anne E Fowler Foundation, Inc.

Disclosed balance after: 5,909

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F2 · ownership.nature: The Reporting Person disclaims beneficial ownership of the shares held by The Anne E. Fowler Foundation, Inc., and this report should not be deemed an admission that the Reporting Person is the beneficial owner of such shares for purposes of Section 16 or for any other purpose.

SEC source: 0001127602-18-028455 · form4.xml · nonDerivative · entry 2

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · by Gillis Family Trust U/a/d 12/30/14

Disclosed balance after: 23,636

Exercise/conversion can include RSU settlement. Transaction price and option strike are different fields; a missing price is not zero. Derivative and underlying rows can describe the same event.

F1 · ownership.nature: Anne Marie Heiser is trustee of the Gillis Family Trust u/a/d 12/30/14.

SEC source: 0001127602-18-027928 · form4.xml · nonDerivative · entry 4

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Gst Exempt Trust - Mk

Disclosed balance after: 0

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.87 to $51.26, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-027928 · form4.xml · nonDerivative · entry 3

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Gst Exempt Trust - Lk

Disclosed balance after: 0

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.87 to $51.26, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-027928 · form4.xml · nonDerivative · entry 2

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Gst Exempt Trust - Dk

Disclosed balance after: 0

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.87 to $51.26, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-027928 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 1,587,657

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.87 to $51.26, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-026453 · form4.xml · nonDerivative · entry 4

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Gst Exempt Trust - Dk

Disclosed balance after: 13,741

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.59, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-026453 · form4.xml · nonDerivative · entry 3

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Gst Exempt Trust - Mk

Disclosed balance after: 13,741

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.59, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-026453 · form4.xml · nonDerivative · entry 2

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Gst Exempt Trust - Lk

Disclosed balance after: 13,741

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.59, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-026453 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 0

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.59, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-026329 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 52,191

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

SEC source: 0001127602-18-026143 · form4.xml · nonDerivative · entry 2

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 52,268

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

SEC source: 0001127602-18-026143 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 52,468

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.08, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-025940 · form4.xml · nonDerivative · entry 2

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 54,773

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F3 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.000 to $50.405, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (3) to this Form 4

SEC source: 0001127602-18-025940 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 205,812

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.22, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-025741 · form4.xml · nonDerivative · entry 2

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 229,738

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F3 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.10, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (3) to this Form 4.

SEC source: 0001127602-18-025741 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 308,736

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

F2 · price: The price reported is a weighted average price. These shares were sold in multiple transactions at prices ranging from $50.00 to $50.02, inclusive. The reporting person undertakes to provide to Athene Holding Ltd., any security holder of Athene Holding Ltd., or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the range set forth in footnote (2) to this Form 4.

SEC source: 0001127602-18-025345 · form4.xml · derivative · entry 2

Sondej John Andrew · CIK 0001298924 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 1,023

Underlying: Class A Common Shares · 1,023 shares

Expires not disclosed

F2 · derivative.exercise_price: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date. An additional 2,046 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2018 to December 31, 2020. Vested RSUs are settled in Class A common shares on a one-for-one basis. In certain circumstances involving termination of the reporting person after a change in control of the Issuer, the award shall become immediately and fully vested as of the effective date of such termination of relationship

F2 · derivative.exercise_date: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date. An additional 2,046 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2018 to December 31, 2020. Vested RSUs are settled in Class A common shares on a one-for-one basis. In certain circumstances involving termination of the reporting person after a change in control of the Issuer, the award shall become immediately and fully vested as of the effective date of such termination of relationship

F2 · derivative.expiration_date: Each restricted stock unit ("RSU") represents a contingent right to receive one Class A common share of the Issuer. The RSUs reported herein vest on each of the first three anniversaries of the January 1, 2018 vesting start date. An additional 2,046 RSUs not reported herein vest only if the Issuer satisfies certain performance criteria over the three fiscal year period from January 1, 2018 to December 31, 2020. Vested RSUs are settled in Class A common shares on a one-for-one basis. In certain circumstances involving termination of the reporting person after a change in control of the Issuer, the award shall become immediately and fully vested as of the effective date of such termination of relationship

SEC source: 0001127602-18-025345 · form4.xml · derivative · entry 1

Sondej John Andrew · CIK 0001298924 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 5,383

Underlying: Class A Common Shares · 5,383 shares

Expires 2028-08-13

F1 · derivative.exercise_date: This option vests ratably on each of the first three anniversaries of the January 1, 2018 vesting start date.

SEC source: 0001127602-18-025221 · form4.xml · nonDerivative · entry 1

Kvalheim Grant · CIK 0001689321 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Indirect · By Grant Kvalheim 2009 Children's Non-Exempt Trust

Disclosed balance after: 311,720

F1 · kind: The sales reported on this Form 4 were effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person on January 9, 2018.

SEC source: 0001127602-18-028455 · form4.xml · nonDerivative · entry 1

Gillis Frank Lauren · CIK 0001689345 · See Remarks

10b5-1 filing: Not disclosed

Ownership: Direct

Disclosed balance after: 289,638